As I understand it, water needs to be oxygenated (replenished)
by churning air (oxygen) with water, otherwise oxygen would
bleed itself out of the water and return to the atmosphere.
So perhaps the water in the harbours are not oxigenated enough
to support the fish ecosystem, they are basically death traps.
On the other hand, the water could be polluted, interfering with
their sensitive bio-systems, also leading to their deaths.
I guess I can understand what is meant by "oxygen deprivation"
and it could take many forms, two as listed above.